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How to know the recipient belongs to which site ?

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there are two sites site-1 and site-2 in a domain, and also there is  a mailbox  named1@domain.com in site-1 and  a mailbox2@domain.com in site-2.

when a user for 1@domain.com in site-1 want to send a mail to2@domain.com ,the hub transport role can get that the2@domain.com is in site-2, so it delivery the mail to the hub transport in site-2, so my question is how did the hub transport know that the2@domain.com located in site-2,?did it get that by inquiry the global catalog,or something else?



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